Once the third stack is added, the enemy champion is struck by lighting after a very brief delay, dealing 30-180 damage, based on level, with AD getting a 40% bonus and AP getting a 25% bonus adaptive damage. Any type of damage, from basic attacks and ability attacks to even item effects and summoner spells, adds a stack. The keystone you will most commonly find when players are primarily using the Domination rune path, Electrocute is a keystone that is built on stacks.
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